
The laws of thermodynamics govern the course of the universe, and all organisms obey these laws. The first law of thermodynamics, also known as the law of conservation of energy, states that energy can neither be created nor destroyed, only transformed or transferred. This law applies to both closed and open systems, and is fundamental to understanding energy transformations and mass distributions in living organisms. Living systems require the creation of order or lower entropy, which may seem to contradict the second law of thermodynamics, but this is possible due to the specific characteristics of open systems. The application of the first law of thermodynamics to life and evolution has been explored by various scientists, including Ludwig Boltzmann, Richard Sears McCulloh, and Erwin Schrödinger, who have contributed to our understanding of the role of energy and entropy in biological processes.
| Characteristics | Values |
|---|---|
| First Law of Thermodynamics | Total energy in a closed system is neither lost nor gained, only transformed |
| Energy can neither be created nor destroyed, only change form | |
| Second Law of Thermodynamics | Entropy constantly increases in a closed system |
| Entropy can decrease or remain constant in an isolated system | |
| Entropy can vary in small-scale systems | |
| Living Organisms | Governed by the laws of thermodynamics |
| Require the temporary creation of order or lower entropy | |
| Can use and store energy | |
| Can use and preserve FGI in their genome | |
| Need to acquire elements, nutrients, and genetic traits | |
| Can extrude end products of metabolism and toxic substances |
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What You'll Learn

The first law of thermodynamics and the origin of life
The first law of thermodynamics, also known as the law of conservation of energy, states that energy can neither be created nor destroyed, only transferred from one form to another. This law applies to all physical systems, including living organisms, which must take in energy from food or other sources and convert it into heat or mechanical work to sustain their physiological functions.
The first law of thermodynamics plays a fundamental role in understanding the origin and evolution of life. It provides a framework for examining the energy transformations and mass distributions that occur within cells and biological systems. For example, cells are semi-permeable, open systems that allow the exchange of mass and energy across their membranes, enabling the acquisition of nutrients and the elimination of waste products.
The origin of life itself can be viewed as a phase transition, governed by the laws of thermodynamics. This transition involves the organization of molecules into more complex structures, eventually leading to the formation of early life forms. While the specific mechanisms remain a subject of ongoing research, scientists have proposed various hypotheses, such as Alexander Oparin's "primordial soup" theory, which suggests that sufficient energy for generating early life forms was provided in a primordial mixture of non-living molecules.
The second law of thermodynamics, which deals with entropy, also plays a crucial role in understanding the origin and development of life. Entropy, as defined by Ludwig Boltzmann, is the struggle for negative entropy or a tendency towards increasing disorder. Living organisms, however, exhibit a decrease or maintenance of entropy, which can be explained by their ability to feed on negative entropy, as suggested by Erwin Schrödinger. This perspective challenges the notion of a naturalistic origin of life, as it implies the need for an external source of information or intelligence to counteract the tendency towards disorder.
In summary, the first law of thermodynamics sets the foundation for understanding the energy exchanges and transformations that occur within living systems. It provides insights into the growth, reproduction, and maintenance of life by examining the flow of energy and matter across cellular membranes. Additionally, the broader framework of thermodynamics, including the second law, helps elucidate the phase transitions and organizational principles that gave rise to the complex structures and functions characteristic of life on Earth.
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The first law and the evolution of life
The first law of thermodynamics states that total energy in a closed system is neither lost nor gained, it is only transformed. This law applies to all particles within a system, governing energy transformations and mass distributions. In other words, energy can neither be created nor destroyed, only changed from one form to another. This is made possible by the fact that cells that comprise living organisms are semi-permeable, open systems that allow both mass and energy to cross their membranes.
The first law has significant implications for the evolution of life. Firstly, it underscores the importance of energy in the evolution of life forms. Living organisms require energy for various processes, including growth, reproduction, and survival. The first law dictates that energy is conserved in these processes, ensuring that the total energy within an organism remains constant. This conservation of energy enables the continuity of life, as energy is passed from one organism to another within an ecosystem.
Additionally, the first law's application to particles within a system highlights its role in the evolution of genetic instructions, or functional genetic instructions (FGI). Genetic variation, resulting from gene transfer and sexual reproduction, leads to increased phenotypic variability and accelerated rates of evolutionary divergence. The first law, by governing energy transformations, influences the expression of genes and the occurrence of genetic errors. When lethal mutations or genetic errors occur, they can disrupt the cell cycle, leading to the death of the organism. This natural selection process ensures that only the fittest FGIs remain and contribute to the evolution of life.
Moreover, the first law's concept of energy transformation is integral to understanding the evolution of life from a thermodynamic perspective. The transformation of energy allows for the creation of order or lower entropy within living systems, which may seem contradictory to the second law of thermodynamics. However, this apparent contradiction is resolved by understanding that living organisms are open systems that exchange matter and energy with their surroundings. This exchange enables life to feed on negative entropy, maintaining and decreasing its own entropy by taking in energy from the environment.
In conclusion, the first law of thermodynamics plays a fundamental role in the evolution of life. It governs energy transformations, influences genetic instructions, and allows for the creation of order within living systems. By conserving energy and facilitating the exchange of matter and energy, the first law provides the framework for the evolution and sustainability of life on Earth.
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Negative entropy and living things
The laws of thermodynamics govern energy transformations and mass distributions. All organisms obey these laws. Living systems cannot persist in isolation. They absorb energy from sunlight or from energy-rich chemical compounds and return part of this energy to the environment as entropy.
Austrian physicist Erwin Schrödinger theorized in his 1944 book, "What is Life?", that life decreases or keeps constant its entropy by feeding on negative entropy. Schrödinger's theory contradicts the second law of thermodynamics, which states that the entropy of an isolated system tends to increase. Schrödinger later corrected himself, stating that the true source is free energy.
The concept of negative entropy, or negentropy, refers to the idea that living things continuously increase entropy. In other words, they create order or lower entropy within their systems, which is made possible by the absorption and transformation of energy.
The struggle for existence is not a struggle for raw materials or energy but for negative entropy, which becomes available through the transition of energy from the hot sun to the cold earth. This transition allows living organisms to acquire the energy necessary to maintain their internal order and counteract the natural tendency towards disorder.
Research concerning the relationship between entropy and the origin and evolution of life has led to various theories and experiments. For example, the alkaline hydrothermal vent (AHV) hypothesis suggests framing life forms as "entropy generators" to understand abiogenesis under alkaline deep-sea conditions. Additionally, the concept of the dissipative system describes the thermodynamics of open systems in non-equilibrium states, providing insights into how living organisms maintain their internal order while interacting with their environment.
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The first law and cellular processes
The laws of thermodynamics are considered the most fundamental laws that govern the workings of the universe. The first law of thermodynamics states that energy in a closed system is neither lost nor gained, it is only transformed. This means that the total energy in the universe or any closed system remains constant.
The first law of thermodynamics is applicable to cellular processes as well. Cells are highly ordered systems with low entropy. They require a constant energy input to maintain this state. This energy is used to perform various cellular processes. For example, in photosynthesis, plants absorb light energy and convert it into chemical energy stored in the form of glucose. This process demonstrates the first law of thermodynamics, as the total energy remains constant, only changing from one form to another.
Another example is cellular respiration, where the energy stored in glucose is released and used by plant and animal organisms to produce ATP. This process again showcases the principle of energy conservation, as the energy from glucose is transformed into a form that can be utilized by the cell.
Furthermore, the first law of thermodynamics is relevant to the activation energy requirement in cells. Most chemical reactions in cells occur too slowly to support the rapid processes necessary for cellular survival. To overcome this, cells use catalysts to facilitate these chemical reactions. By providing catalysts, cells can lower the activation energy barrier, enabling reactions to occur more readily without violating the first law, as the energy is conserved throughout.
The first law of thermodynamics, therefore, plays a crucial role in understanding the fundamental workings of cellular processes. It highlights the importance of energy conservation and transformation in maintaining the highly ordered state of cells and facilitating the various processes essential for their survival.
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The first law and the growth cycle
The first law of thermodynamics is a formulation of the law of conservation of energy in the context of thermodynamic processes. It states that energy cannot be created or destroyed, only transformed from one form to another. This law is fundamental as it governs the course of the universe, and all organisms, including life, obey it.
Living organisms require nourishment to survive and flourish, and this is where the first law of thermodynamics comes into play. The first law and the concept of growth are closely intertwined. The law of conservation states that potential energy is always finite for any entity, be it an individual, a family, or a business. However, the first law also tells us that energy can be transformed from one form to another. This means that resources can be strategically invested and utilized to generate growth. For instance, a company requires revenue growth to achieve its vision and ensure its longevity. Similarly, individuals need nourishment to sustain themselves and grow.
In the context of life, the first law of thermodynamics is closely linked to the growth cycle. Cells, which comprise living organisms, are semi-permeable, open systems that allow both mass and energy transfer across their membranes. This enables the acquisition of essential elements and nutrients, as well as the extrusion of metabolic end products and toxic substances. Cell division is accompanied by reproduction, and death can occur either as a programmed or accidental consequence. The first law guides cells through the cell cycle, and genetic instructions (FGI) play a crucial role in this process.
The first law of thermodynamics also has implications for the origin of life and its further development. While there is no standard model for how the first biological life forms emerged, the laws of thermodynamics impose certain constraints on the earliest life-sustaining reactions. To remain consistent with the second law of thermodynamics, self-organizing systems with lower entropy values than equilibrium must dissipate energy to increase entropy in their external environment. This relationship between the first and second laws of thermodynamics influences the growth and reproduction of life, as well as the battle against entropy or disorder.
In summary, the first law of thermodynamics, with its principle of energy conservation, underpins the growth cycle of life. It dictates the acquisition and transformation of energy, influencing the survival, growth, and reproduction of living organisms. The law's interplay with the second law of thermodynamics shapes the evolution and ongoing battle against entropy, ensuring the fitness and continuity of life.
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Frequently asked questions
The first law of thermodynamics states that total energy in a closed system is neither lost nor gained, only transformed. In other words, energy can neither be created nor destroyed, only changed from one form to another.
Yes, life does obey the first law of thermodynamics. Living organisms are open systems that allow mass and energy to cross their membranes. This means they can acquire elements and nutrients and expel metabolic waste and toxins. All organisms obey the laws of thermodynamics, which govern energy transformations and mass distributions.
Entropy is a measure of randomness or disorder in a system. Living things continuously increase entropy, which is known as "negative entropy" or "negentropy". This means that life adds to the overall entropy of the universe. However, life also requires the temporary creation of order or lower entropy, which seems to contradict the second law of thermodynamics.
The second law states that entropy constantly increases in a closed system. This presents a challenge for understanding the origin of life, as there is no "standard model" of how the first biological life forms emerged. The second law implies that self-organizing systems with lower entropy than equilibrium must dissipate energy to increase entropy in their external environment.










































